Lordkipanidze et. al. published the paper A complete skull from Dmanisi, Georgia, reveals evolutionary biology of early Homo, in Science, saying that this is the only complete fossil Homo skull of the early Pleistocene ever found so far. It provides direct evidence for wide variation within populations of early Homo, implying a single evolving lineage with continuity across continents.
The site of Dmanisi, Georgia, has yielded an impressive sample of hominid cranial and postcranial remains, documenting the presence of Homo outside Africa around 1.8 million years ago. Here we report on a new cranium from Dmanisi (D4500) that, together with its mandible (D2600), represents the world's first completely preserved adult hominid skull from the early Pleistocene. D4500/D2600 combines a small braincase (546 cubic centimeters) with a large prognathic face and exhibits close morphological affinities with the earliest known Homo fossils from Africa. The Dmanisi sample, which now comprises five crania, provides direct evidence for wide morphological variation within and among early Homo paleodemes. This implies the existence of a single evolving lineage of earlyHomo, with phylogeographic continuity across continents.